MORPHOLOGY
External characteristics
Male
Forewings: The forewing upper side has a large black border. The forewing underside is black coloured. The is no colouring in the bottom-most cell space on the anal margin (Pequin, 2021). Hindwings The hindwing underside has never white coloured veins around the cell (Pequin, 2021).
Female
The females generally have a different underside wing surface than the males, by being darker with broader margins on both wings (D'Abrera, 1982).
Subspecies and variation
Four subspecies are described, of which two subspecies are found in Papua and two subspecies are found in Irian Jaya. The ssp hiemalis has a form, f. hitam, of which the forewing underside is totally black coloured. Ssp. flabella has a pale subterminal area with short orange brown streaks. The forewing upper side of ssp. nemangkawi has a reduced white area (Pequin, 2021).